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The cell's eating habits in the Cell Stage directly influence its diet in the Creature Stage, and only mouths appropriate to the diet (Herbivore, Carnivore, or Omnivore) established in the Cell Stage will become available in the Creature Stage.

Each phase has three characteristics, usually based on how aggressively or peacefully the phase was played.

Characteristics determine how the creature will start the next phase and give it abilities that can be used later in the game.

Once the primary objective is completed, the player has the option to advance to the next stage, or to continue playing the current stage.

The Cell Stage (sometimes referred to as the tide pool, cellular, or microbial stage) is the very first stage in the game, and begins with a cinematic explanation of how the player's cell got onto the planet through the scientific concept of panspermia, with a meteor crashing into the ocean of a planet and breaking apart, revealing a single-celled organism.

Phases often feature optional missions; when the player completes a mission, they are granted a bonus, such as a new ability or money.

If all of a player's creations are completely destroyed at some point, the species will be respawned at its nearest colony or at the beginning of the phase.

These are then automatically uploaded to the online Sporepedia and are accessible by other players for download.